Bacon said:
Try disconnecting the feedback, and cut your gain way back and try some more. If the problem remains, it will probably be worse, and that would mean it's not the feedback but maybe a bad transformer, a bad driver tube etc. But if the problem basically goes away, then it is the feedback.
I will admit that I never worked on one but I have to agree with Bacon on this one. It sounds like some kind of impedance unbalance in the driving circuit. Instead of swapping drivers, swap the leads on the driving Xformer if possible. If you wind up with 100% positive vs. 50% negative, then you know you found the problem. Gud Luck.