That's pretty much what I said on the first page of this thread; apparently, you missed it.
Actually, I had tried adjusting the screen voltage previously as you had suggested and it didn't make a noticeable difference. However this was before the installation of the 15H choke on the output of the screen voltage power supply and the realization that the 4-400As would not work well at low power which is the power level that I was doing most of my earlier measurements at. After eliiminating other variables like the output network tuning, I recalled that I hadn't tried adjusting the PA screen voltage since I had installed the 15H choke in series with the screen voltage dropping resistor.
Once I get the audio feedback issues sorted out, I anticipate that I will be adjusting all of the operating parameters for best performance.
I very much appreciate everyone's help and suggestions!